V. R. Alifanova, S. V. Savelievb, E. Yu. Tereshchenkoc, V. V. Artemovc, and A. Yu. Sereginc
aBorissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya ul. 123, Moscow, 117997 Russia
e-mail: valifan@paleo.ru
bScientific Research Institute of Human Morphology, Russian Academy of Medical Sciences,
ul. Tsyurupy 3, Moscow, 117418 Russia
e-mail: embrains@hotmail.com
cShubnikov Institute of Crystallography, Russian Academy of Sciences, Leninskii pr. 59, Moscow, 117333 Russia
e-mail: helena.tereshenko@ gmail.ras.ru; artemov@ ns.crgs.ras.ru
Received July 23, 2013
Abstract—Integumentary structures of ornithischain dinosaurs of the taxon Hypsilophodontia (Ornithopoda)
from the Ukureiskaya Formation (Upper Jurassic) of the Kulinda locality (Transbaikal Region, Russia) are
described in detail. It is shown that members of this group had so-called bristle scales, integumentary append-
ages previously unknown in ornithischian dinosaurs. These are relatively small horn plates embedded in the
skin, the distal margin of which has several long, flat, and probably constantly growing bristles. The monobris-
tle variant of bristle scale is probably homologous to the protofeather of theropods; if this is the case, it is pos-
sible to reconstruct the protofeather as an elongated and constantly growing scale.
Keywords: Dinosauria, Ornithischia, Ornithopoda, Hypsilophodontia, skin, skin derivatives, bristle scales
DOI: 10.1134/S0031030114050025
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